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It is obvious that even the problem of _coadaptation in sterile animals_ can thus be satisfactorily explained. If the determinants are oscillating upwards and downwards in continual fluctuation, and varying more pronouncedly now in one direction now in the other, useful variations of every determinant will continually present themselves anew, and may, in the course of generations, be combined with one another in various ways. But there is one character of the determinants that greatly facilitates this complex process of selection, that, after a certain limit has been reached, they go on varying in the same direction. From this it follows that development along a path once struck out may proceed without the continual intervention of personal selection. This factor only operates, so to speak, at the beginning, when it selects the determinants which are varying in the right direction, and again at the end, when it is necessary to put a check upon further variation. In addition to this, enormously long periods have been available for all these adaptations, as the very gradual transition stages between females and workers in many species plainly show, and thus this process of transformation loses the marvellous and mysterious character that seemed at the first glance to invest it, and takes rank, without any straining, among the other processes of selection. It seems to me that, from the facts that sterile animal forms can adapt themselves to new vital functions, their superfluous parts degenerate, and the parts more used adapt themselves in an ascending direction, those less used in a descending direction, we must draw the conclusion that harmonious adaptation here comes about _without the cooeperation of the Lamarckian principle_. This conclusion once established, however, we have no reason to refer the thousands of cases of harmonious adaptation, which occur in exactly the same way among other animals or plants, to a principle, the _active intervention of which in the transformation of species is nowhere proved. We do not require it to explain the facts, and therefore we must not assume it._

For example, a very extraordinary story is told of one of these Saxon princesses. A certain king upon the Continent, whose dominions lay between the Rhine and the German Ocean, had proposed for her hand in behalf of his son, whose name was Radiger. The consent of the princess was given, and the contract closed. The king himself soon afterward died, but before he died he changed his mind in respect to the marriage of his son. It seems that he had himself married a second wife, the daughter of a king of the Franks, a powerful continental people; and as, in consequence of his own approaching death, his son would come unexpectedly into possession of the throne, and would need immediately all the support which a powerful alliance could give him, he recommended to him to give up the Saxon princess, and connect himself, instead, with the Franks, as he himself had done. The prince entered into these views; his father died, and he immediately afterward married his father's youthful widow--his own step-mother--a union which, however monstrous it would be regarded in our day, seems not to have been considered any thing very extraordinary then.

• Nicholas Bredimus
• Nicholas Bredimus
• Nicholas Bredimus
• Nicholas Bredimus
• Nicholas Bredimus

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